DNA alignment of complete gag-pol frame of MuERVC-C105, MuRRS, MuRVY and active viruses MoMLV and GALV.

The defective MuERV sequences have numerous in-frame terminators and frameshifts. In some places, the alignment can not be objectively defended and reasonable alternative alignments would trade frameshfits for in-frame terminators. Regions of poor alignment among MuERV sequences correspond to regions of poor conservation among active retroviruses. Regions of relatively secure alignment used for phylogenetic reconstructions are indicated.

Numbers refer to base under final digit and match numbering system in the GenBank files defined in overview. For the numbering of MuERVC, see MuERVC-C105 sequence It is AF049340 with the 718 bp L1C105 insert removed starting at coordinate 180. First letters of gag-pol codons have been capitalized. All 3 letters of in-frame terminators have been capitalized.

SCH 6/3/98


                    630       640       650       660       670       680
     MoMLV   AtgGgcCagActGttAccActCccTtaAgtTtgAccTtaGgtCacTggAaaGatGtcGag
             ||||| ||   |  | | || ||  |     | ||  ||  |||||||| ||||||
     GALV    AtgGgaCaaGatAatTctAccCctAtcTccCtcActCtaAatCacTggAgaGatGtgAga
                    640       650       660       670       680       690
             |||||| |   |    | |||||  |  | || ||||  || || || | ||| || |
     MuRVY   AtgGgaAagActGtgAcgAccCcaCtaAcaCttActCagAacCatTGAAcaGacGttAtg
            1080      1090      1100      1110      1120      1130
             |  |||  |||||| |||||||| ||||   | |||    |||| || || || ||||
     MuRRS   ActGgaCggActGtaAcgAccCccCtaAgtTtgActTtaGacCacTggActGaaGttAga
           1250      1260      1270      1280      1290      1300

    MuERVC   >>>>>>>>>>>>>>>>>>>>>>>>>>>>> This portion not cloned >>>>>>
             Met  --> start of gag  MA domain


                       690       700       710       720       730
     MoMLV   CggAtc...GctCacAacCagTcgGtaGatGtcAagAagAgaCgtTggGttAccTtcTgc
                |     |||||||| |  || || ||  |||| ||| ||   |||   || |||||
     GALV    AcaAgg...GctCacAatCtaTccGtgGaaAtcAaaAagGgaAaaTggCagActTtcTgt
                       700       710       720       730       740
             || |||   ||  ||||||| | | | ||  |||| ||||||   ||||| ||||| ||
     MuRVY   ActAgg..gGccAacAatCttTtcTttGatGtcAagAagGgaCctTggCaaActTttTgc
            1140        1150      1160      1170      1180      1190
              | |||   ||| | || ||||   | ||  | |||||||| |||||||| ||||| |||
     MuRRS   TcaAgg...GccCatAacCttTcaGtaGaaAttAagAagGggCctTggCagActTtcTgc
           1310         1320      1330      1340      1350      1360

    MuERVC   >>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>
         


             740       750       760       770       780       790
     MoMLV   TctGcaGaaTggCcaAccTttAacGtcGgaTggCcgCgaGacGgcAccTttAacCgaGac
             ||  | || ||||| || ||   ||| || ||||| |  || || || ||||| |
     GALV    TccTccGagTggCccAcaTtcGgcGtgGggTggCcaCcgGagGgaActTttAatCtcTct
             750       760       770       780       790       800
             |||||  | |||||  | || || ||||| |||||  || |||||| |||||  ||  ||
     MuRVY   TccTctAaaTggCcaGctTttGgtGtgGgcTggCctTcgCagGgaAatTttAgcCtgCct
              1200      1210      1220      1230      1240      1250
              ||||  | | |||  |||||  |||    ||| |   | | || | |||| |  |  ||
     MuRRS   GccTccGagTAGCcgActTttAatGtaAaaTggTcaCtgGaaGggActTtt.ggTttAct
              1370      1380      1390      1400      1410       1420

    MuERVC   >>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>
         


             800       810       820       830       840       850
     MoMLV   CtcAtcAccCagGttAagAtcAagGtcTttTcaCctGgcCcgCatGgaCacCcaGacCag
              ||||       ||||| |  |  ||||||        |      ||||| || |||||
     GALV    GtcAttTttGcaGttAaaAagAttGtcTttCagGagAac...GggGgaCatCcgGacCaa
             810       820       830       840          850       860
             || ||| | |  || ||     || | ||||||         || || ||||| || |
     MuRVY   GttAttCtaGgtGtcAa....GttAttTttCagActGca...GgaGggCatCctGatCcg
              1260      1270          1280      1290         1300
              ||||  | |  || ||     || ||||||||| || |   ||    || |  ||||
     MuRRS   CttAtcTttGaaGttAaaGccAttGttTttCagAgtGgaCctGggTccCacCtgGatCaa
               1430      1440      1450      1460      1470      1480

    MuERVC   >>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>
         


             860       870       880       890       900       910
     MoMLV   GtcCccTacAtcGtgAccTggGaaGccTtgGctTttGacCccCctCccTggGtcAagCcc
             || || || ||||||   ||| | | | | ||     | ||||| || |||||     ||
     GALV    GttCcaTatAtcGtgGtaTggCagGacCtcGccCagAatCccCcaCcaTggGtgCcaGcc
                870       880       890       900       910       920
             || || |||||  | || |||||||| || |  || ||||  ||||| ||  ||    ||
     MuRVY   GtaCctTatAtaAttGtgTggCagGatCtgGtaCaaAatCtgCcaCccTGACtgGgtTcc
           1310      1320      1330      1340      1350      1360
                |||||                   ||  | || |||   || || | | |     |
     MuRRS   CagCctTac..................CtcAttCagAatTcaCcgCcaTAAAtcAaaCca
               1490                        1500      1510      1520

    MuERVC   >>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>
                                  End of portion used as MA-><----------


             920       930                   940       950       960
     MoMLV   TttGtaCacCctAagCctCcg............CctCctCttCctCcaTccGccCcgTct
             |                                  |     |  | |
     GALV    Tcc..................GccAagGtcGctGttGtcTctGatAccCga.........
                                  930       940       950
             |                    || | ||| || || |  |  || |||  |
     MuRVY   TggActTccGgaGctGagGcaGcgAtgGtgGcaGtaGcaT..GaaAccGaaAatGat...
           1370      1380      1390      1400        1410      1420
             |||  |
     MuRRS   TggGtt......................................................
             1530

    MuERVC   >>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>
             --- alignment considered unreliable in this region


               970       980       990      1000            1010
     MoMLV   CtcCccCttGaaCctCctCgtTcgAccCcgCctCgaTccTcc......CttTatCcaGcc
                ||  |||        |  ||  | || || ||| || ||       | || || ||
     GALV    AgaCcaGttGcgGggAggCcaTcaGctCctCccCgaCccCcc......AtcTacCcgGca
             960       970       980       990            1000      1010
             ||||||   | |      |      |    |  || |||||       ||||| || | |
     MuRVY   AgaCcaTgcGggTcaGccCagAgcAcgTtcCatCggCccCctGagG..AtcTatCctGaa
                1430      1440      1450      1460        1470      1480
                ||||||   ||||  |    | |          |   |
     MuRRS   CccCcaTgcCtcTcaGagCccTccCccGgaTgcTccCatGca..................
                 1540      1550      1560      1570

    MuERVC   >>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>GAtcTatCcaGag
                                                            1       10


                                     1020      1030      1040      1050
     MoMLV   ...........................CtcActCctTctCtaGgcGccAaaCctAaaCct
         
     GALV    AcaGacGacTta................................................
                  1020
             |  || ||
     MuRVY   AttGatGatGcc................................................
                  1490

     MuRRS   ............CtcGagTggGtcAgcAcaGtgAccTcgCttTctGcc...CtaAaa.ag
                           1580      1590      1600      1610
                         |  || |||     |||
    MuERVC   AtaGaaGaaCctCcaGaaTggTgtTacAcc..............................
                  20        30        40


                 1060      1070      1080      1090      1100      1110
     MoMLV   CaaGttCttTctGacAgtGggGggCcgCtcAtcGacCtaCttAcaGaaGacCccCcgCct
                                        ||| ||  |       |       ||||||||
     GALV    ...........................CtcCtcCtcTctGaaCccAcg...CccCcgCcc
                                           1030      1040         1050
                                         || |   |||             || || ||
     MuRVY   ........................TttTtcAttGacTccTtc.........CcaCccCct
                                        1500      1510               1520
                                         |   |      ||
     MuRRS   CagTccCtg...............CacCtgGctCtgAgaTtgCta...............
           1620                     1630      1640

    MuERVC   ...................................................CcaCccCaa
                                                                     50


                 1120      1130      1140      1150      1160      1170
     MoMLV   TatAggGacCcaAgaCcaCccCctTccGacAggGacGgaAatGgtGgaGaaGcgAccCct
             |||  ||   ||   |||||||||   | |    | |  |  ||        |   ||
     GALV    TatCcgGcgGcaCtgCcaCccCctCtgGccCctCagGcgAtcGgaCcgCcgTcaGgcCag
                1060      1070      1080      1090      1100      1110
             || ||      |    |    |||    || | |||  |  |   ||  |      |  |
     MuRVY   TacCccCaaAgaActTctAatCctGatCccTcaCagCtgCccAccCcaGcaGacCccAgg
                   1530      1540      1550      1560      1570      1580

     MuRRS   ............................................................
         

    MuERVC   TatCccCaaCtgAtcCccTccCagAtcTctCaaCagGgtGctGctGgtGatGcgGaa...
                   60        70        80        90       100


                 1180      1190      1200      1210      1220
     MoMLV   GcgGgaGagGcaCcgGacCccTccCcaAtgGcaTctCgcCtaCgtGggAgaCgg...Gag
               |   ||       || ||          ||  || |     |  |  | ||
     GALV    AtgCccGatAgtAgcGatCctGagGggCcaGccGctGggAccAggAgtCgcCgtGccCgc
                1120      1130      1140      1150      1160      1170
              || ||  |               || ||    |  |||||    ||  ||   | ||
     MuRVY   GtgGccTctGtg.........AcaGgaCcgTtgGgaGggActCctAgcTgcGaaGtcCat
                   1590               1600      1610      1620      1630
                    |   |           ||||||    |  ||||| |  ||| |  ||| |||
     MuRRS   ...AagGcaCagTctGagGacCgaGgaCctAcaGctGggAccCggAgcCgaCaaGccCag
              1650      1660      1670      1680      1690      1700
                                     |||||  | ||||| | ||  ||| |   ||| |||
    MuERVC   .....................GagGgaCcaGctGctGgaAacCaaAgcTggAgaGctCag
                                110       120       130       140


          1230      1240                        1250      1260      1270
     MoMLV   CccCctGtgGcc..................GacTccActAccTcgCagGcaTtcCccCtc
                || |  | |                  |||||||||        |   || ||||||
     GALV    AgtCcaGcaGac......AacTcgGgtCctGacTccAct......GtgAttTtgCccCtc
                1180            1190      1200            1210      1220
             ||| |||           || ||    || || ||||||      ||   ||||||  |
     MuRVY   AgtTcaGagAggGaaGtgAa.TctAaaCcaGatTccAct......GttGctTtgCctTta
                  1640      1650       1660            1670      1680
             ||| |              |       |  ||    ||       |    ||| |||||
     MuRRS   AgtCccCcaGca......GatGaaGctCttGac...AccActTctGcc..tTttCctTtg
              1710            1720      1730         1740        1750
             || ||                   |  |  |||   |||       ||    | |||||
    MuERVC   AgcCcaGag...............GgaCcaGacTcaAcc......AccGagCtaCctTta
            150                      160       170             180


                  1280                           1290               1300
     MoMLV   CgcGcaGga......GgaAac...............GgaCagCtt.........CaaTac
             || ||   |      |||  |                     ||          |||||
     GALV    CgaGccAta......GgaCcc...CcgGccGagCccAatGgcCtgGtcCctCtaCaaTat
                  1230               1240      1250      1260      1270
              ||||          |||||    ||       |         |  ||||  | || ||
     MuRVY   AgaGctTatGtaGgaGgaCc..tgCccAtgCccCggGg.CagTtaAtcCcaTtgCagTac
               1690      1700        1710      1720       1730      1740
              |||||         ||     || |||    | ||       |     |  |  | ||
     MuRRS   TGAGctGtg......GgcAacCtgTccAcaGacAggCagTctCtgCagAccCtcTAATat
               1760            1770      1780      1790      1800
                 ||         |||   |   |  |||||     |    ||||| ||||  ||||
    MuERVC   CatActTat......GgcCctCctCcaGcaGacCccGggCaaTtgCagCccCttCaaTac
             190             200       210       220       230       240
                        alignment considered unreliable -------------><--


                  1310                  1320      1330      1340
     MoMLV   TggCcgTtcTccTct............TctGacCttTacAacTggAaaAatAatAacCct
             ||||| || |||||              | || ||||| || ||||||  |||| | ||
     GALV    TggCctTttTccTca............GcaGatCttTatAatTggAaaTctAatCatCcc
                 1280                  1290      1300      1310      1320
             ||||| ||||| |              |||||| | |||||           ||||  ||
     MuRVY   TggCcaTttTcaTtgAtaGatAtgTtgGcaGatAtgTatAac........aTatCacTcc
                  1750      1760      1770      1780              1790
             ||||| |||||||              || ||| |||| ||
     MuRRS   TggCctTttTcaTca............GctGatCtgTacAatTAG...............
           1810      1820                  1830      1840
             || |||||||| |||             ||||||| || ||||||
    MuERVC   TgtCctTttTccTca............TctGatCtcTatAatTAGAagAcaAatCatGcc
                   250                   260       270       280
             ---  Region used as CA domain

          1350      1360      1370      1380      1390      1400
     MoMLV   TctTttTctGaaGatCcaGgtAaaCtgAcaGctCtgAtcGagTctGttCtcAtcAccCat
             |||||||||||| | ||||     || || |  ||  | |||||| || |  || |||||
     GALV    TctTttTctGaaAacCcaGcaGgtCtcAcgGggCtcCttGagTctCttAtgTtcTccCat
                   1330      1340      1350      1360      1370      1380
             ||||| ||  | || ||  ||||  | || || ||||||||||  || ||||||||| |
     MuRVY   TctTtcTcaTAGAatCccTcaGgcTtaAcaGgaCtcCttGagTtcCtaAtgTtcTccTac
                1800      1810      1820      1830      1840      1850

     MuRRS   ............................................................
         

    MuERVC   GccTtcTct.....cTccTcaAgcCttActAatCtaAtaGaaTctCtcAtgTtcTctCat
           290            300       310       320       330       340


          1410      1420      1430      1440      1450      1460
     MoMLV   CagCccAccTggGacGacTgtCagCagCtgTtgGggActCtgCtgAccGgaGaaGaaAaa
             |||||||| |||||||| || || |||||  |   || |||  | |||   || |||
     GALV    CagCccActTggGacGatTgcCaaCagCtcCtaCagAttCttTtcAccActGagGaaCgg
                   1390      1400      1410      1420      1430      1440
             |||||||||||||| || ||||| || ||  | ||| | |  || |  ||||| ||  |
     MuRVY   CagCccActTggGatGacTgcCagCaaCttTtgCagGtcCccTttAgaActGaaGagAga
                1860      1870      1880      1890      1900      1910

     MuRRS   ............................................................
         

    MuERVC   CaaCctActTggGatGacTgtCagCagCttTtaCagGttCttTtcAccAcaGggGagAag
                350       360       370       380       390       400


          1470      1480      1490         1500      1510      1520
     MoMLV   CaaCggGtgCtcTtaGagGctAga...AagGcgGtgCggGgcGatGatGggCgcCccAct
              || |  | ||| | |||||  |    ||    || |  || ||  |||||  ||| ||
     GALV    GaaAgaAttCtcCtgGagGccCgc...AaaAatGtcCttGggGacAatGggGccCctAca
                   1450      1460         1470      1480      1490
             || || || || || || ||| |    ||||| ||||  ||     ||||| |||| ||
     MuRVY   GagAggAtcCtgCtaGaaGccAgaA..AaaAacGtcCcaGgaCcaGatGggAccCccAct
                1920      1930        1940      1950      1960      1970

     MuRRS   ............................................................
         

    MuERVC   GaaAagAttTggCtgGagGcaAga...AaaAatGtgCtaGggGctAatAgaCtaCtgAca
                410       420          430       440       450       460


             1530      1540      1550      1560      1570      1580
     MoMLV   CaaCtgCccAatGaaGtcGatGccGctTttCccCtcGagCgcCcaGacTggGatTacAcc
             || ||    ||     |  |||  || || |||||| | || ||  |||||||||||| |
     GALV    CagCtcGagAacCtcAttAatGagGccTtcCccCtcAatCgaCctCacTggGatTacAac
            1500      1510      1520      1530      1540      1550
              | |||   ||||| || ||||  || |||||| |    ||  ||  ||  || | |||
     MuRVY   AatCtcCccAacCttAtaAatGcaGctTtcCccTtgGccCgtTctGccTtaGagTtcAat
                  1980      1990      2000      2010      2020      2030
                               || || |  |||||  ||| |||  | |  |   | | |||
     MuRRS   ..................AacGctGggTtcCctCtgGacCggCcaGatTAGAacTacAac
                                    1850      1860      1870      1880
                                | || || |||||||||    | || || ||| ||   |||
    MuERVC   CaaGttCtgAatGaaAtaGatGcaGgaTtcCctCtgAcaTgtCctGacTAGGacCctAac
                   470       480       490       500       510       520


             1590      1600      1610      1620         1630      1640
     MoMLV   AccCagGcaGgtAggAacCacCtaGtcCacTatCgc...CagTtgCtcCtaGcgGgtCtc
             ||    ||||||||| | |  ||  |   ||| |||   | |   ||  | || ||||||
     GALV    AcaGccGcaGgtAggGagCgtCttCtgGtcTacCgc...CggActCtaGtgGcaGgtCtc
            1560      1570      1580      1590         1600      1610
             || |    || ||||||||||||    |||||| |     ||||||||||||||||  |
     MuRVY   ActGtgAaaGatAggGagCgtCtcAcaGtcTacTGA...TggActCtaGtgGcaGggAta
                  2040      2050      2060         2070      2080
             || |   ||| || |||| |  | ||||| |||||     || |||||||        ||
     MuRRS   AcaGcaCaaGgtAagGagAggTtgAcaGttTacTgg..cCggGctCtaGt.......Cta
                1890      1900      1910        1920      1930
             |   |  |||||| ||||  ||||||| ||| | |    |||||||| ||       ||
    MuERVC   AggCctGaaGgtAggGagCagTtgAcaAttTtcCgc...CggGctCtgGtaGcaGgaCtg
                   530       540       550          560       570


                1650      1660      1670      1680      1690      1700
     MoMLV   CaaAacGcgGgcAgaAgcCccAccAatTtgGccAagGtaAaaGgaAtaAcaCaaGggCcc
              ||   || |   |  | || ||||||||||| ||||||| ||   |    || || ||
     GALV    AaaGggGcaGctCggCgtCctAccAatTtgGctAagGtaAgaGagGtcTtgCagGgaCcg
               1620      1630      1640      1650      1660      1670
              |  |||| ||  || | || |||||||||||  |||||||||| ||| | |||||  |
     MuRVY   GagCggGctGcaTggTGACccAccAatTtgGccTAGGtaAgaGaaGtcCttCagGgtTca
           2090      2100      2110      2120      2130      2140
              || | ||     |  | ||||||||||||||  |||||||||||||  | |||||
     MuRRS   AagGgaGcgAtgAgaCgcCccAccAatTtgGctAagGtaAgaGaaGtgAtgCagGgcCtt
               1940      1950      1960      1970      1980      1990
             |  |||||    ||  ||||||||| |||||| |||||||||||  |  | || ||  |
    MuERVC   AgaGgaGctGgcAggTgcCccAccAgtTtgGccAagGtaAgaGagAtcCtaCaaGggTta
            580       590       600       610       620       630


                1710      1720      1730      1740      1750      1760
     MoMLV   AatGagTctCccTcgGccTtcCtaGagAgaCttAagGaaGccTatCgcAggTacActCct
                ||  | || ||||  ||| ||||  | || | ||| |||||| | || ||||||||
     GALV    GcaGaaCccCctTcgGttTtcTtaGaaCgcCtgAtgGagGccTatAggAgaTacActCcg
               1680      1690      1700      1710      1720      1730
                || |  || ||  ||||||||| ||| || |||||| | |||||||||||    ||
     MuRVY   AtgGagCtgCccTccAttTtcTtaGcaCgaCtaAtgGagTctTatAggAgaTa..gcCca
           2150      2160      2170      2180      2190      2200
             |  ||||  || ||  |||| |||| | || | || ||  |||||||    ||     |
     MuRRS   AcaGagCccCcaTctGttTttTtaGaaAgaTtgAtaGaaGctTatAgaTacTatActTct
               2000      2010      2020      2030      2040      2050
             |  || |||| | | || | ||||||        |||| || || ||  | || ||| |
    MuERVC   AagGaaCccCtaAcaGtgTgtTtaGagCacCccTtaGagGcaTacAggCatTacActCcc
            640       650       660       670       680       690


                1770      1780      1790      1800      1810         1820
     MoMLV   TatGacCctGagGacCcaGggCaaGaaActAatGtgTctAtgTctTtc...AttTggCag
             | ||| ||          || ||| |  ||   ||  | ||| | ||    ||  | |||
     GALV    TttGatCccTctTctGagGgaCaaCagGctGcgGtcGccAtgGccTtt...AtcGgaCag
               1740      1750      1760      1770      1780         1790
             |||||||| ||||     ||||| |||      || ||||| |  ||    || || |||
     MuRVY   TttGatCc.TctT....aGgaCagCagC....tGtaGccAtaGatTta..cAttGgtCag
             2210           2220          2230      2240        2250
             ||| | ||  | |     ||  | ||       || || || | |||
     MuRRS   TttAacCcgAccTctGagGgtTAACaaGccGcgGttGctAtgGctTtt............
               2060      2070      2080      2090      2100
             ||| |||| |  || ||| |  |||| |   | || || |
    MuERVC   TttGacCctAtaTcaGagAgcCaaCagGtgAcaGtaGccA..............TggCag
            700       710       720       730                     740


                         1830      1840      1850      1860      1870
     MoMLV   TctGccCca......GacAttGggAgaAagTtaGagAggTtaGaaGatTtaAaaAacAag
             || ||||||      || ||   || ||||||| ||||| |||| |   |  |  || |
     GALV    TcaGccCca......GatAtcAagAaaAagTtaCagAggCtaGagGggCtcCagGacTat
                  1800            1810      1820      1830      1840
             || | | |       ||||| | || ||||||||||||| ||||||| ||||| || |||
     MuRVY   TctGacTct......GatAttAtgAgaAagTtaCagAggTtaGagGgtCtcCaaGatTat
              2260            2270      2280      2290      2300
                                                        |  |  | || |
     MuRRS   ...........................................aaAgaTtaCatGccAtg
                                                            2110
                                                        |  |  | ||  |  |
    MuERVC   TctGctTctGacAt.GacAtcAggAgaAggTtaCatTgtCtaGagGgtCtgCaaCcaTta
                750        760       770       780       790       800


               1880      1890         1900      1910      1920
     MoMLV   AcgCttGgaGatTtgGttAgaGag...GcaGaaAagAtcTttAatAaaCgaGaa...Acc
              |  |   |||||| || | ||||   ||||||||| | |   ||||  ||||    ||
     GALV    TccTtaCaaGatTtaGtaAaaGag...GcaGaaAagGtgTacCatAagAgaGag...Aca
              1850      1860         1870      1880      1890
              |  | ||| |||||||||||||    || || || |||||||| |||||||||   ||
     MuRVY   ActCtgCaaCatTtaGtaAaaGaaA..GctGagAaaGtgTacCacAagAgaGag...Act
          2310      2320      2330        2340      2350      2360
              || | ||| ||||||| || ||    ||  |||| ||||| | |||| ||||    |
     MuRRS   GctTtaCaaGatTtaGttAagGag...GcaCagAagGtgTatCccAagGgaGaaT..Aag
          2120      2130      2140         2150      2160      2170
              |||| |||||| | |||||||||   ||| |||| ||||| |  ||| ||||    |
    MuERVC   TctTtgCaaGatCtgGttAagGag...GcaGagAaaGtgTacCatAagAgaGag...Act
                 810       820          830       840       850
                                       End region used as CA ------------

           1930      1940            1950      1960      1970      1980
     MoMLV   CcgGaaGaaAga......GagGaaCgtAtcAggAgaGaaAcaGagGaaAaaGaaGaaCgc
                |||||||||       | ||| |    |  |  ||  |||| ||||| ||    ||
     GALV    GaaGaaGaaAga......CaaGaaAgaGaaAaaAagGagGcaGaaGaaAagGagAggCgg
          1900      1910            1920      1930      1940      1950
             || |||||||        ||||| |||||| | || |||| ||| ||||  || |  |||
     MuRVY   GagGaaGaaAag......CaaGagAgaGaaGagAaaGagGtaGagGaaAgaGaaAacCgg
               2370            2380      2390      2400      2410
               ||||  ||
     MuRRS   AggGaaAgaAga................................................
               2180
                 ||  |  |
    MuERVC   GaaAaaTAAGaaAaaA..CgaGaaAgaGaaAagAgaGagGctGagGcaAgaGaaAgcAga
             860       870         880       890       900       910
             -----------><--- Alignment not used through to pol domain
                              although pro region has some usable segments. 

                 1990      2000      2010      2020      2030      2040
     MoMLV   CgtAggAcaGagGatGagCagAaaGagAaaGaaAgaGatCgtAggAgaCatAgaGagAtg
                                               | ||| |   ||   | | | |  ||
     GALV    .................................CgcGatAggCcgAagAaaAaaAacTtg
                                                 1960      1970      1980
                                              |  ||  | | | || | |  ||||||
     MuRVY   .................................CatGaaCgcCagGagGatAggAacTtg
                                            2420      2430      2440
                                                           |||| ||||| ||
     MuRRS   .............................................CagGagAggAatTta
                                                          2190      2200
                                                          || || || ||||||
    MuERVC   .................................TGAGatAgaAggCaaGaaAgaAatTta
                                                920       930       940


                 2050      2060                        2070      2080
     MoMLV   AgcAagCtaTtgGccActGtcGttAgt..................GgaCagAaaCag...
             |  ||  |  ||||| | || || |||                   || |
     GALV    ActAaaAttCtgGccGcaGtaGtaAgt..................AgaGaaGggTccAca
                   1990      2000                        2010      2020
             ||||| ||| |||| ||||| ||| ||                   ||||| | ||
     MuRVY   ActAagAttTtgGcgGcaGtgGtaGgt..................GgaGaaAgtTcaGag
              2450      2460      2470                        2480
             |  | |||||||||  |||| |||||                    |  |  |  |||||
     MuRRS   AgcAggAttTtgGccTcaGtaGtaGggGaaTgtAaaGaa......AggAacGggCcaGag
                  2210      2220      2230      2240            2250
             |  | |||||| |   || |   ||| ||    || |||      || || ||   ||
    MuERVC   ActAagAttTtaGatAcaTtcAcaGgtGat...AagGaaGttTatAgaAaaGgtAgaGga
                  950       960       970          980       990


                  2090                                    2100
     MoMLV   GatAgaCagGgaGga..............................GaaCgaAggAgg...
             | ||| |||  |||                                      | ||
     GALV    GgtAggCagAcaGggAacCtgAgcAacCagGcaAagAagAcaCctAggGatGgaAga...
                 2030      2040      2050      2060      2070
                 | ||| ||||| ||||| |||||  |||||  |   | ||  | ||    |||
     MuRVY   AagGgtCagTcaGggTacCtgCgcAacAggGcaAgaAgaCccCcaGgaGaaCagAga...
            2490      2500      2510      2520      2530      2540
             ||  |  || ||||||||||| |||||| || |   |||   ||   |  |   |||
     MuRRS   AatAgaTAGAcaGggTacCtgGgcAacAagGaaCccAga...CctAaaAgaAgaAgaTct
              2260      2270      2280      2290         2300      2310
             | |||  |||||||||||||||||||||  | ||  |||   ||| | |     |||
    MuERVC   AgtAggCagAcaGggTacCtgGgcAacAcaGgaCtgAgaAgaCctCagAacTccAgaGaa
          1000      1010      1020      1030      1040      1050


                    2110      2120      2130      2140      2150
     MoMLV   .........TccCaaCtcGatCgcGacCagTgtGccTacTgcAaaGaaAagGggCacTgg
                       | | ||| ||    |||||||| || ||||| ||||| ||||| || |||
     GALV    .........CctCcaCtaGacAaaGacCagTgcGcaTacTgtAaaGagAagGgcCatTgg
                   2080      2090      2100      2110      2120      2130
                      |||||||| || |||||||| ||| ||||||| ||  | || |   |||||
     MuRVY   .........CctCcaCttGagAaaGacCacTgcAcaTacTgcAagAaaAaaGaaTatTgg
                        2550      2560      2570      2580      2590
                             | ||||| ||||| ||  | || |||||| |||||| | || ||
     MuRRS   ...GtcCagAagAttTtaGagAagGacCaaTgtGccTatTgcAagGaaAaaGgaCatAgg
                    2320      2330      2340      2350      2360
                   ||      | | || |||||||||||||| |  | ||| ||||| |||||  ||
    MuERVC   CctTcaCaaCcaCctCtgGatAagGacCaaTgtGcaTgcTacAaaGaaAagGgaCacTgg
          1060      1070      1080      1090      1100      1110


          2160      2170      2180      2190      2200      2210
     MoMLV   GctAaaGatTgtCccAagAaaCcaCgaGgaCctCggGgaCcaAgaCccCagAccTccCtc
             || | ||| ||||||   |||  |         |  |    |  | || ||        |
     GALV    GcaAgaGaaTgtCccCgaAaaAaa.........CacGtcAgaGaaGccAagGttCtaGcc
                   2140      2150               2160      2170      2180
             || ||||| || ||   |||              |    || |   | |||  |||  ||
     MuRVY   GccAgaGacTgcCcaAaaAag............AagCagAggGgcCcaAagTatCtgCcc
              2600      2610                  2620      2630      2640
             ||  |||| |||||||| ||             |||   |||| ||| |||   ||| |
     MuRRS   GcaCgaGaaTgcCcaAagAaaA...........AagGgaAggGccCctAagGtcCtgAct
          2370      2380      2390                 2400      2410
             ||| |||| ||||| |  |||             |    ||  | || |||||||| ||
    MuERVC   GcaAgaGacTgcCccAgaAaaGagAca......GatAccAgaCcaCccAagGtcCttAcc
          1120      1130      1140            1150      1160      1170


          2220      2230         2240      2250      2260      2270
     MoMLV   CtgAccCtaGatGacTAG...GgaGgtCagGgtCagGagCccCccCctGaaCccAggAta
                   |||||| |||||   ||  ||||||||  ||| |||| ||| ||||| ||| ||
     GALV    ......CtaGatAacTAG...GggAgtCagGgtTcgGacCccCtcCccGaaCctAggGta
                        2190         2200      2210      2220      2230
                     |||  || ||   || |||| ||| |||| |||||||||| | |||||||||
     MuRVY   Tt....GaaGaaGacAag...GgaAgtCggGgcTcgGtcCccCtcCccAagCctAggGta
                    2650         2660      2670      2680      2690
              |     |||| || ||     | || |||||||| |  ||||||||| |||||||||||
     MuRRS   Ctg...AaaGatGatAaa...TggAgaCggGgcTcaGggCccCtcCccGagCctAggGta
           2420         2430         2440      2450      2460      2470
             ||     | ||| |  |     ||||| |||||||||  ||||||||| || ||||||||
    MuERVC   CtaGaaGatGatAacTAG...GggAgaTggGgcTcaGacCccCtcCccAagTctAggGta
                1180      1190         1200      1210      1220      1230
                        Gag Ter

             2280      2290      2300         2310      2320      2330
     MoMLV   AccCtcAaaGtcGggGggCaaCcc...GtcAccTtcCtgGtaGatActGggGccCaaCac
             || || |  || | ||||   |||    |    |||||||| || || || ||  ||||
     GALV    AcaCtgActGtgGagGggAccCcc...AttGagTtcCtgGtcGacAccGgaGctGaaCat
                 2240      2250         2260      2270      2280
             || |   |||||||||||   |||    || | || ||  | ||||| ||||||||| ||
     MuRVY   ActCacTctGtgGagGggTtaCccCc.GttAatTttCtaAtaGacAcaGgaGctGaaTat
             2700      2710      2720       2730      2740      2750
             || |     ||| | | |   ||     |  ||||| |||| |||||||| || ||| ||
     MuRRS   AccCtgAtgGtgAaaGagActCca...AtgGatTttTtaAttGacAcaGggGcaGaaCat
                 2480      2490         2500      2510      2520
             ||||| |  ||| | | ||| ||    | ||||||  |  |||| || || |||||   |
    MuERVC   AccCtaActGtgGagGggAccCcc...AcgGatTtcCttGttGatAccGgaGcaGagTtt
                   1240      1250         1260      1270      1280


                2340      2350      2360      2370      2380      2390
     MoMLV   TccGtgCtgAccCaaAatCctGgaCccCtaAgtGatAagTctGccTggGtcCaaGggGct
             || ||  ||||||||      |||    || |    |      |    ||  |||| ||
     GALV    TcaGtaTtgAccCaaCccAtgGgaAaaGtaGggTccAgaCggAcgGtcGtgGaaGgaGcg
          2290      2300      2310      2320      2330      2340
             ||||| |||||  ||||  | ||| || |||| || |    ||   | ||      |||
     MuRVY   TcaGtcTtgActGaaCctTtaGgaCaaTtaGgaTcaAagAagAttTtaGttAtcCcaGcc
              2760      2770      2780      2790      2800      2810
             || || ||||   | ||  |  || |||||  |   || |  |   |||| ||   ||||
     MuRRS   TctGtgTtgAaaCatCcaCtgAgaAaaTtaAaaAatAaaAgaAccAtaGtgAttGgaGcc
          2530      2540      2550      2560      2570      2580
             || ||  ||||| | |||||| | ||||| |  || || | |||  |||| || || ||
    MuERVC   TcaGtcCtgAaaGagCcaCtgGggAaaTtgAggAacAagAaaActGtaGtaAtcGgcGct
            1290      1300      1310      1320      1330      1340


                2400      2410      2420      2430      2440      2450
     MoMLV   ActGgaGgaAagCggTatCgcTggAccAcgGatCgcAaaGtaCatCtaGctAccGgtAag
             || ||  | |||   || | |||||||||  |  |     || |  | |         |
     GALV    AcaGgcAgcAagGtcTacCccTggAccAcaAaaAgaCttTtaAaaAttGgaCatAaaCaa
          2350      2360      2370      2380      2390      2400
             || || |||||| | || |||||||| || |||||  ||||| | || ||| | ||  ||
     MuRVY   ActGgtAgcAagCttTatCccTggAcaAccAaaAggAttTtaGacAtaGgaAaaAatAaa
              2820      2830      2840      2850      2860      2870
             ||||||   || |  || |||||||| ||   | | | | ||||  | || |||    ||
     MuRRS   ActGgtCagAaaCaaTacCccTggAccActGcaTgcActGtaGatCtgGggAaaGgcCaa
          2590      2600      2610      2620      2630      2640
             |||||||||||||||||  | ||||||||||   | |   | ||| |||| | | |||||
    MuERVC   ActGgtCagAaaCaaTatTctTggAccActGatAggAaaCttGatTtgGgaAgaAgcCaa
            1350      1360      1370      1380      1390      1400


                2460      2470      2480      2490      2500      2510
     MoMLV   GtcAccCacTctTtcCtcCatGtaCcaGacTgtCccTatCctCtgTtaGgaAgaGatTtg
             || |||||||| |||||     |||| || || ||   ||||||||| || || ||  |
     GALV    GtgAccCacTccTtcCtgGtcAtaCccGagTgcCctGctCctCtgTtgGgcAggGacCtc
          2410      2420      2430      2440      2450      2460
             ||||| || || || || || ||||| |||||||| || || ||| | ||| |||||||
     MuRVY   GtgAcgCatTctTttCttGtgAtaCctGagTgcCcaGcaCccCtgCtaGgcTggGacCtt
              2880      2890      2900      2910      2920      2930
             ||||  ||||| ||  | || || |||||||| ||  |||| || |||||  | || ||
     MuRRS   GtgAgtCatTccTtcTtgGtcAttCctGagTgtCccAcaCcgCttCtaGgaAgaGatCtg
          2650      2660      2670      2680      2690      2700
             ||   ||||| |||| | ||||||||||| |||||||| ||  |  |||| |||||  |
    MuERVC   GtaGctCatTacTtcCtcGtcAttCctGaaTgtCccAccCcaTtgTtaGggAgaGacTta
            1410      1420      1430      1440      1450      1460


                2520      2530      2540      2550      2560      2570
     MoMLV   CtgActAaaCtaAaaGccCaaAtcCacTttGagGgaTcaGgaGctCagGttAtgGgaCca
             || || |||||||| ||||| ||||| |||   |     ||  | ||||| |   |   |
     GALV    CtaAccAaaCtaAagGccCagAtcCagTttTccGctGagGgcCcaCagGtaAcaTggGga
          2470      2480      2490      2500      2510      2520
                |  ||| | || || ||  |||||||| |  |||| || ||  ||||| |||||
     MuRVY   TgtAaaAaaTtgAaaGcaCaaGtcCagTttActTctGatGgaCctGagGtaTcaTggAag
              2940      2950      2960      2970      2980      2990
                |  ||||| || || ||  ||   ||||| |  || || ||    ||| | ||| |
     MuRRS   ...ActAaaTtaAagGccCagAtcGgaTttAccTAAGaaGggCcgAgaGtaActTggGaa
             2710      2720      2730      2740      2750      2760
                 | |||||||| ||||| ||    ||  | |  |||||  |   |||  ||||  ||
    MuERVC   CtaTcaAaaTtaAaaGccCaaAtaAacTtcTctTtgGaaGgaActTcaGtcTctTGACaa
            1470      1480      1490      1500      1510      1520


                2580      2590      2600      2610      2620      2630
     MoMLV   AtgGggCagCccCtgCaaGtgTtgAccCtaAatAtaGaaGatGagCatCggCtaCatGag
                   |  ||           ||  ||||||  | ||||| ||  | || ||||||||
     GALV    Gaa...CgcCctActAtgTgcCtgGtcCtaAacCtgGaaGaaGaaTacCgaCtaCatGaa
          2530         2540      2550      2560      2570      2580
              |    |           ||| | ||  ||||  | ||||||||||| |  || || |||
     MuRVY   AagAatCca...GtgGcaTgcTtaGttTtaAatTtaGaaGaaGaaTatCacCtcCacGaa
              3000         3010      3020      3030      3040      3050
                   ||       | |  |||||   |  |  | || |||||||| |   | || |||
     MuRRS   Tct...CccAccTccGtaGtcTtaGccCtgCagCttGagGaaGaaTacCgaTtgCatGaa
                2770      2780      2790      2800      2810      2820
                   || | |||  || | |||||  |  |  | || ||||||||  |||| |||||
    MuERVC   Gtc...CctAtcTctAtaAttTtaGctTtaAacTtaGaaGaaGaaTatAgaTtaCatGag
            1530         1540      1550      1560      1570      1580


                2640      2650      2660      2670      2680      2690
     MoMLV   AccTcaAaaGagCcaGatGttTctCtaGggTccAcaTggCtgTctGatTttCctCagGcc
             |   ||  |            ||| | |   |  | |||||      ||| ||    |
     GALV    AagCcaGtaCcc......TccTctAtcGacCcaTccTggCtcCagCttTtcCccActGta
             2590            2600      2610      2620      2630      2640
              ||                  |    |     |    |  |      ||| ||    ||
     MuRVY   CagAagTctGagAct......TtaCccTttAaaGaaCgaTtgAcaGgtTttCcaGagGtt
                 3060            3070      3080      3090      3100
                |      | ||         |  | | |||||  | ||||| | ||| ||    |
     MuRRS   GgcAtaAagCaaAcaGagGtgCcaGacCtaAaaGacTggTtgActGctTtcCctAgaGca
                2830      2840      2850      2860      2870      2880
                  |   | | ||        |     |     ||||||       |  |||  |||
    MuERVC   CctCaaGccCcaTcaAgaAaaGtaCtaGaaGggAccTggTtaGagAgaTatCctCaaGct
               1590      1600      1610      1620      1630      1640


                2700      2710      2720      2730      2740      2750
     MoMLV   TggGcgGaaAccGggGgcAtgGgaCtgGcaGttCgcCaaGctCctCtgAtcAtaCctCtg
             ||||| ||||  |  ||||||||||| ||   ||      | ||  || |  ||   ||
     GALV    TggGcaGaaAgaGccGgcAtgGgaCtaGccAatCaaGtcCcaCcaGtgGtaGtaGagCta
                   2650      2660      2670      2680      2690      2700
             ||||||||    || || |||| | |||| |  ||||| |||||     |||| ||||||
     MuRVY   TggGcaGagCagGctGggAtgGaaTtaGctAggCaaGtgCcaCccAacAtaGtcGagCta
             3110      3120      3130      3140      3150      3160
             |||||||||   || || |||||| | |||     ||| || ||     | || |  ||
     MuRRS   TggGcaGagActGcaGgcAtgGaaAtgGctGtcAgaGttCccCctGtgGttGtgGgaCtg
                2890      2900      2910      2920      2930      2940
             ||||||||||||| ||| |||  |  |||    || || ||| |  |  |||||   |||
    MuERVC   TggGcaGagActGgaGgtAtgAgaGagGcaAagAggGtgCccTccAtcAttGtgAtgCtg
               1650      1660      1670      1680      1690      1700
                                                       <- Begin pol domain

                2760      2770      2780      2790      2800      2810
     MoMLV   AaaGcaAccTctAccCccGtgTccAtaAaaCaaTacCccAtgTcaCaaGaaGccAgaCtg
             | | ||    |  | || ||| |  |   |||||| || |||    |||||||  |
     GALV    AgaTcaGgtGccTcaCcaGtgGctGttCgaCaaTatCcaAtgAgcAaaGaaGctCggGaa
                   2710      2720      2730      2740      2750      2760
             | |   | ||||  |||  |  | ||  | ||||| || |||||  |||  |||    |
     MuRVY   AaaAtgGctGccAaaCccAttTcaGtcAggCaaTacCccAtgAgtCaaGggGctAaaAac
             3170      3180      3190      3200      3210      3220
             || | || ||| |  || ||   |||    ||||| || ||||| |     || | | |
     MuRRS   AagAcgGatGcaAccCctAtaGgaGtgCaaCaaTatCcaAtgAgcCgtAaaGcaAgaGat
                2950      2960      2970      2980      2990      3000
             || || | ||  || |||||  |  |   |||||| || |||| |    | ||   |||
    MuERVC   AaaActGgtGtgActCctAtcTgg.taTGACaaTacCccAtgAacAaaGagGccCaaGag
               1710      1720       1730      1740      1750      1760


                2820      2830      2840      2850      2860      2870
     MoMLV   GggAtcAagCccCacAtaCagAgaCtgTtgGacCagGgaAtaCtgGtaCccTgcCagTcc
             || ||||  |||||||| ||||   |  | ||||  ||  |  |||| ||||| | |||
     GALV    GgtAtcAgaCccCacAtcCagAagTtcCtaGacCtaGggGtcTtgGtgCccTgtCggTcg
                   2770      2780      2790      2800      2810      2820
              | ||  |||| || |||||||   | ||| | |  ||  | ||||| || || | |||
     MuRVY   AgcAttCgaCcaCatAtcCagAgaCtgCtaTAACagGgtAtaTtgGtcCcgTGACagTcc
             3230      3240      3250      3260      3270      3280
              | ||  | |||||||| ||||||||  || ||   ||||| ||||| || || |||||
     MuRRS   GgaAtcAggCcaCatAttCagAgaCtaTtaCaaTtaGgtAttTtgGtaCctTgcCagTca
                3010      3020      3030      3040      3050      3060
             || ||||| || || ||||| ||     |    |  | | |||||||    |  |||||
    MuERVC   GgcAtcAgaCctCacAttCaaAg....GtgAggTctGatGttTtgGtcAt.TatCagTct
                1770      1780          1790      1800       1810


                2880      2890      2900      2910      2920      2930
     MoMLV   CccTggAacAcgCccCtgCtaCccGttAagAaaCcaGggActAatGatTatAggCctGtc
             |||||||| || || |||||||| || || || |||||||| ||||| ||| |||| ||
     GALV    CccTggAatAccCctCtgCtaCctGtaAaaAagCcaGggAccAatGacTatCggCcaGtt
                   2830      2840      2850      2860      2870      2880
             |  |||||||||||  | || || ||    || || ||||| |||||||||  ||| ||
     MuRVY   CatTggAatAccCccTtcCtgCcaGtgTgtAaaCctGggAcaAatGacTatGtgCctGtc
             3290      3300      3310      3320      3330      3340
             |  |||||||| || || ||| | ||    |||||||| || | ||||||    || |||
     MuRRS   CccTggAatActCctTtaCtgTcgGtaAagAaaCctGgcAcgAgtGacTacTGACcaGtc
                3070      3080      3090      3100      3110      3120
             |||||||||||||| || ||  | || || || || ||||| | ||| ||  | |||||
    MuERVC   CccTggAatActCcaTtgCtaCcaGttAaaAagCcaGgcActAatGatTatCggCcaGta
           1820      1830      1840      1850      1860      1870


                2940      2950         2960      2970      2980      2990
     MoMLV   CagGatCtgAgaGaaGtc...AacAagCggGtgGaaGacAtcCacCccAccGtgCccAac
             || || ||||||||| |    || ||  ||||  | || || || ||||| || || |||
     GALV    CaaGacCtgAgaGaaAtt...AatAaaAggGtaCagGatAttCatCccAcaGtcCcaAac
                   2890         2900      2910      2920      2930
             || ||| | ||| || |    |||||  |||||||||| |||||  | || || || |||
     MuRVY   CagGacTtaAgaAaaGtcC..AatAagCggGtaCagGacAttCaaTcaActGtgCccAac
             3350      3360        3370      3380      3390      3400
             || ||| | ||| | |||   |||||| |||| || || |||||  ||||||| || ||
     MuRRS   CaaGacCtgAgaGagGtc...AatAagAggGtcCaaGatAttCacCcaActGttCctAat
                3130      3140         3150      3160      3170      3180
             |||||| | ||||| ||    || || || || ||||||||||| || ||||| |||||
    MuERVC   CaaGacTtaAgaGaaGta...AacAaaAgaGtaCaaGatAttCatCctActGtcCctAac
           1880      1890         1900      1910      1920      1930


                   3000      3010      3020      3030      3040      3050
     MoMLV   CctTacAacCtcTtgAgcGggCtcCcaCcgTccCacCagTggTacActGtgCttGatTta
             |||||||| ||  ||||    || || ||   | |    |||||| | ||  | ||| |
     GALV    CctTacAatCttCtgAgtTccCttCcgCctAgcTatActTggTacTcaGtcTtaGatCtc
            2940      2950      2960      2970      2980      2990
             ||| ||||  | ||||||||| | || |||       | |||||| | ||| | ||| |
     MuRVY   CctCacAacTtgCtgAgtTccTtaCcaCctGaaCggTccTggTacActGtcCtgGatTtg
               3410      3420      3430      3440      3450      3460
             ||  | ||| | || || ||  | || || || |     ||||| || || || |||||
     MuRRS   CccTatAacCtcCtcAgcTcaCtcCctCcgGagCaaAaaTggTatAcaGtaCttGatTta
                   3190      3200      3210      3220      3230      3240
             ||||| |  || ||||| || ||||| || ||   | |||||||||  ||||| ||  |
    MuERVC   CccTacA.tCtaCtcAgtTctCtcCccCctGacAgaCaaTggTatAtgGtaCtgGacCtg
              1940       1950      1960      1970      1980      1990


                   3060      3070      3080      3090      3100      3110
     MoMLV   AagGatGccTttTtcTgcCtgAgaCtcCacCccAccAgtCagCctCtcTtcGccTttGag
             |||||||||||||||||||| || || || |||| ||| ||||| || ||||| || |||
     GALV    AagGatGccTttTtcTgcCtcAggCtaCatCccAacAgcCagCcgCtgTtcGcgTtcGag
            3000      3010      3020      3030      3040      3050
             || |||||||||||||| |||||  |||| |  ||||| || |  ||||  || || |||
     MuRVY   AaaGatGccTttTtcTgtCtcAgaTtaCacCtgAacAgtCaaCttCtgTgtGctTttGag
               3470      3480      3490      3500      3510      3520
             |||||||| || || || || |  ||||| |    ||| || |   | |  || ||||||
     MuRRS   AaaGatGctTtcTttTgcCtgAagTtaCatCccTccAgcCagCccAtaTtcGccTttGag
                   3250      3260      3270      3280      3290      3300
             || ||  |  |||| ||||| ||  ||||||||   ||   |   || || || ||||||
    MuERVC   AagGacAc..tcTtcTgcCtaAaaCtaCatCccAtgAgtTggGtaAtcTttGctTttGag
               2000        2010      2020      2030      2040      2050


                   3120      3130      3140      3150      3160
     MoMLV   TggAgaGatCcaGagAtgGgaAtcTcaGgaCaaTtgAccTggAccAgaCtcCcaCag...
             |||| ||| ||||| |  || | | |||| ||  ||||||||||  | || |||||
     GALV    TggAaaGacCcaGaaAaaGgtAacAcaGgtCagCtgAccTggAcgCggCtaCcaCaa...
            3060      3070      3080      3090      3100      3110
             |||    |||| |||  |||  | || |||||||| |  | |||  |||| | |||
     MuRVY   TggGggTacCctGaaGgaGgaTatAcgGgtCagCtaAttTtgActAggCtgCgaCagA..
               3530      3540      3550      3560      3570      3580
             ||| |  | |||||    ||| | ||||| ||  | |  | ||   ||   |  |||
     MuRRS   TggAgaGatCctGacActGgaCaaAcgGgaCaaTtgAccTggAtgTggTAACccCag...
                   3310      3320      3330      3340      3350
             |||||||| || ||  | |||||  | |||||| | || ||||     |  || ||
    MuERVC   TggAgaGacCcaGatTcgGgaCagGctGgaCaaCtaActTggAcaCatTtgCctCaa...
                 2060      2070      2080      2090      2100


            3170      3180      3190      3200      3210      3220
     MoMLV   GgtTtcAaaAacAgtCccAccCtgTttGatGagGcaCtgCacAgaGacCtaGcaGacTtc
             || ||||| |||  |||||| || || || ||||| || ||| ||||  | ||   |||
     GALV    GggTtcAagAacTctCccActCtcTtcGacGagGccCtcCacCgaGatTtgGctCccTtt
               3120      3130      3140      3150      3160      3170
             || ||||| || || |||||| | || || || |   ||||||   |  |     |||||
     MuRVY   GgaTtcAaaAatTcaCccActTttTttGatGaaGtgGtcCacCacAacCttAtgTccTtt
                 3590      3600      3610      3620      3630      3640
             |  |||||||| || |||||  |||||||||| |   | || |   |  |    |||||
     MuRRS   GagTtcAaaAacTccCccAccCttTttGatGagGctCtaCatCgaGatTtaGccTccTtc
            3360      3370      3380      3390      3400      3410
             |  ||||| ||||| || || || ||||||||||| || ||  | || || ||    |||
    MuERVC   GgaTtcAagAacTctCctActCtcTttGatGagGccCtcCacTggGacTtgGcaCagTtc
          2110      2120      2130      2140      2150      2160


            3230      3240      3250      3260      3270      3280
     MoMLV   CggAtcCagCacCcaGacTtgAtcCtgCtaCagTacGtgGatGacTtaCtgCtgGccGcc
              ||  ||   ||||  |  || |  | || || || ||||| ||| |  || ||||||||
     GALV    AggGccCtcAacCccCagGtgGtgTtaCtcCaaTatGtgGacGacCtcTtgGtgGccGcc
               3180      3190      3200      3210      3220      3230
             ||||  |  |||| |||  |     | ||  | ||||  || ||  |  |  | || ||
     MuRVY   AggGaaCagAacCtcCaaAttTctCtcCtgGagTatGgaGatGatTtaAtcCttGcaGct
                 3650      3660      3670      3680      3690      3700
              | |   | |||  |||  |  |||| ||  | ||||  ||||| ||  ||||||| ||
     MuRRS   CgaGctGaaAacTccCagGtaActCtgCttCaaTatGttGatGacTtgCtcCttGctGca
            3420      3430      3440      3450      3460      3470
              |  | |||||| ||||  |  |||| ||||| ||||||||||||  ||| ||||||||
    MuERVC   AgtTcaGaaAacCccCaaAttTctCtcCttCagTatGttGatGacCggCttCttGctGcc
          2170      2180      2190      2200      2210      2220


            3290      3300      3310      3320      3330      3340
     MoMLV   ActTctGagCtaGacTgcCaaCaaGgtActCggGccCtgTtaCaaAccCtaGggAacCtc
              |  |  |   ||||||| || |||| || | |   || |||||     || | ||  |
     GALV    CccAcaTatGaaGacTgcAaaAaaGgaAcaCagAagCtcTtaCagGagTtaAgtAagTtg
               3240      3250      3260      3270      3280      3290
              | ||  | |||   |||    |  | |   |||||||  || | ||   |||| | |||
     MuRVY   TcaAccCagGaaCtgTgcCttGacAggAtcAagAagCttCtaAatGaaAaaAgtGaaTtg
                 3710      3720      3730      3740      3750      3760
              | ||  |   |   |||    |  | | |||||   | |||   |||    |||| |||
     MuRRS   AccAcgGa.AgaGaaTgcTggCagGgaAccAagAgaTtgCtaGcaGaaCtgGgtGagTtg
            3480       3490      3500      3510      3520      3530
             |||||  |   ||| ||| | |  ||| |  ||| ||| || |||||| || | || |||
    MuERVC   AccAccAagCaaGagTgcAgaCtaGgaCctGagAaaTtaCtgGcaGaaTtgAgcGaaTtg
          2230      2240      2250      2260      2270      2280


            3350      3360      3370      3380      3390      3400
     MoMLV   GggTatCggGccTcgGccAagAaaGccCaaAttTgcCagAaaCagGtcAagTatCtgGgg
             ||||| ||||  ||||| ||||| |||||  | ||||||| | | ||||  |||||||||
     GALV    GggTacCggGtaTcgGctAagAagGccCagCtcTgcCagAgaGaaGtcAccTatCtgGgg
               3300      3310      3320      3330      3340      3350
             | ||||          ||||||| || ||||| ||||| |  ||  | ||||| || ||
     MuRVY   GtgTac........tCctAagAaaGctCagCtaTgcCatAtgGacTtgAccTacCtaGga
                         3770      3780      3790      3800      3810
             |  |||        | |||| ||||| |||||||| || |||||  ||  ||||||  ||
     MuRRS   GgtTacCggGccTctGctAaaAaaGccCagCtaTgtCagAtgGaaGtgGtcTacCtgAga
             3540      3550      3560      3570      3580      3590
              | |||  |||  |  |||| |||| |||| | || ||||  |||||   |||| || |
    MuERVC   AgaTacAagGcaCccActAatAaaGtcCagAttTgcCagAcaGaaGtaAccTacTtgGgt
          2290      2300      2310      2320      2330      2340


            3410      3420      3430      3440      3450      3460
     MoMLV   TatCttCtaAaaGagGgtCagAgaTggCtgActGagGccAgaAaaGagActGtgAtgGgg
             || || || || || ||  | |||||||| ||    ||| |||| |  ||||| |||
     GALV    TacCtaCtcAagGaaGgaAaaAgaTggCtaAccCcaGccCgaAagGctActGttAtgAaa
               3360      3370      3380      3390      3400      3410
             |||   |||   ||||||||    ||| |  ||  |||  |||||   ||||| ||| |
     MuRVY   TacTccCtcCgaGaaGgaAagCagTggTtcTccGaaGctGgaAagAagActGtgAtgCag
               3820      3830      3840      3850      3860      3870
             ||  | |  || || | ||| | |||| |  | || ||| |||| | ||||||||  ||
     MuRRS   TatActC..CggGatGaaAaaCggTggCtgAcaGagGctAgaAaaAggActGtgActCaa
             3600        3610      3620      3630      3640      3650
             |||||||   |||  |  |||| |||||||||||| |||   |||| |||||||||||||
    MuERVC   TatActCtcAggGgaGggAaaCagTggCtgAcaGaaGctCagAaaAagActGtgActCaa
          2350      2360      2370      2380      2390      2400


            3470      3480      3490      3500      3510      3520
     MoMLV   CagCctActCcgAagAccCctCgaCaaCtaAggGagTtcCtaGggAcgGcaGgcTtcTgt
                |||  ||| | ||||||  ||||  |  | || || ||||| || || || |||||
     GALV    AtcCctGttCctAcgAccCccAgaCagGtcCgtGaaTttCtaGgcActGccGgaTtcTgc
               3420      3430      3440      3450      3460      3470
             |||||    || || || |   |||| || ||||| ||||| || |  || || || ||
     MuRVY   AtcCcaAccCccAccActCtgTGACaaGtaCgtGagTttCtgGgaAagGcaGgcTttTgt
               3880      3890      3900      3910      3920      3930
             || ||||| ||  | ||||   |||| ||  | || ||  |||| |  || || || ||
     MuRRS   AtaCcaActCcgGctActCcaAgaCagGtgAgaGaaTtcTtgGgcActGctGggTtcTgc
               3660      3670      3680      3690      3700      3710
             || || |  ||  | ||  | |  |||||||| |||||  ||||||  |||||||| ||
    MuERVC   AtcCccAtcCcaAcaAccTcgAagCagGtgAggGaaTttCtgGgcAtgGctGggTttTgt
          2410      2420      2430      2440      2450      2460


            3530      3540      3550      3560      3570      3580
     MoMLV   CgcCtcTggAtcCctGggTttGcaGaaAtgGcaGccCccTtgTacCctCtcAccAaaAcg
              | ||||||||||||||||||||     |||| || ||||||||||| || || |||  |
     GALV    AggCtcTggAtcCctGggTttGctTccCtgGctGcaCccTtgTacCccCtaAcaAaaGag
               3480      3490      3500      3510      3520      3530
             || ||||| || || | ||   |     |||| |||||  ||||||| || || |||||
     MuRVY   AgaCtcTGAAtaCcaGagTccAcaAtgTtgGcgGcaCctCtgTacCcaCtcActAaaGaa
               3940      3950      3960      3970      3980      3990
             ||||| || ||||| | ||  ||||   |||| || ||     |   ||| || || |||
     MuRRS   AgaCttTggAtaCctGggTttAcaAcaCtgGctGccCcc...CatGtaCtgAccAagGaa
               3720      3730      3740      3750         3760      3770
             || || ||||| ||  | ||| | || ||||| |||||     |    || || || ||
    MuERVC   AggCtcTggAtcCcaAgcTttGccActCtgGcaGccCcaCtcTacCccCttActAaaGag
          2470      2480      2490      2500      2510      2520


            3590      3600      3610      3620      3630      3640
     MoMLV   GggActCtgTttAatTggGgcCcaGacCaaCaaAagGccTatCaaGaaAtcAagCaaGct
              | |  |  |||| ||||      || || ||  |||| | | |  | || ||  ||||
     GALV    AgcAtcCctTttAttTggActGagGaaCatCagCagGctTttGacCacAtaAaaAaaGcc
               3540      3550      3560      3570      3580      3590
             |   |||| || |  ||||| |||||||| ||  | || ||| ||  || |||    ||
     MuRVY   AaaGtcCcgTtcAccTggAcaGagGaaCacCaaAaaGccTttAacAgcAcaAagGctGct
               4000      4010      4020      4030      4040      4050
             |  |     || |                                       |||  ||
     MuRRS   AgtGgaGaaTttAga.....................................agGagGcc
                  3780                                           3790
             | ||     ||   |                                     |  |||||
    MuERVC   ActGcgActTtcTcaTggAccCccAtaCacCaaGagGccTttAaaGagAttAaaAagGcc
          2530      2540      2550      2560      2570      2580


            3650      3660      3670      3680      3690      3700
     MoMLV   CttCtaActGccCcaGccCtgGggTtgCcaGatTtgActAagCccTttGaaCtcTttGtc
              | ||  | ||||| ||  |||   | |||||  | || ||||| ||    || | | |
     GALV    TtgCtgTcaGccCctGcaTtgGccCtcCcaGacCtcAccAagCcaTtcActCtaTatAta
               3600      3610      3620      3630      3640      3650
              ||||  || |||  || |||||||| |||||  | || ||||| |||||  ||||  |
     MuRVY   CtgCttGcaAccCagGctTtgGccCtgCcaGatTtgActAagCctTtcAccAtaTacGtt
               4060      4070      4080      4090      4100      4110
             |||||  |  | |  || ||||| ||||||||| | || ||||| |||| | | || |||
     MuRRS   CtgCtgActGcgCcaGc.TtgGcaCtgCcaGatCtaAccAagCccTtcAtcCtcTatGtt
               3800      3810       3820      3830      3840      3850
              |||| | ||| |||||  | || || |  || || || ||||| ||||  || |||||
    MuERVC   TtgCttAgtGcaCcaGcaCtaGccCttCtgGacCtgActAagCctTtcActCtaTatGta
          2590      2600      2610      2620      2630      2640
                End pol----><---Begin RNAse domain

            3710      3720      3730      3740      3750      3760
     MoMLV   GacGagAagCagGgcTacGccAaaGgtGtcCtaAcgCaaAaaCtgGgaCctTggCgtCgg
             || ||||     |||   |||   || || || || || |   | ||||| ||||| |||
     GALV    GatGagAgaGccGgcGtgGccCggGgaGtgCtcActCagActTtaGgaCccTggCggCgg
               3660      3670      3680      3690      3700      3710
             ||||||  ||| || || |||||  |||| || ||||||||| | || ||||||  | |
     MuRVY   GatGagCcaGctGgaGtaGccCgcAgaGtcCtgActCagActCtgGggCccTggAagAg.
               4120      4130      4140      4150      4160      4170
             ||||||   || || || ||| | ||||| ||  ||||| |||| ||||| ||| ||||
     MuRRS   GatGagAggGcgGggGtgGccAgaAgaGttCttTctCagGctCtaGggCcgTggGagAgg
                3860      3870      3880      3890      3900      3910
             ||  | || || ||  | || ||| |  | ||  | || ||||| ||||| | | |  |
    MuERVC   GacAaaAgtGcaGgaAtaGcgAgaGggTtcCtgAccCaaGctCtgGggCcaTAGAaaCga
          2650      2660      2670      2680      2690      2700


            3770      3780      3790      3800      3810      3820
     MoMLV   CcgGtgGccTacCtgTccAaaAagCtaGacCcaGtaGcaGctGggTggCccCctTgcCta
             || || || || || || ||||| || || || || ||    ||||||||  | |||||
     GALV    CcaGtaGcaTatCtaTcaAaaAaaCtgGatCcgGtgGccAgcGggTggCcaAccTgcCtg
               3720      3730      3740      3750      3760      3770
             ||||| || || || || |||||| |||| || || ||||| |  |||||  |||||||
     MuRVY   CcaGtgGccTacCtgTctAaaAaaTtgGacCcaGttGccAgtGaaTggCccTccTgcCta
                4180      4190      4200      4210      4220      4230
             |  ||||| ||  ||||||| ||  |||||||||| ||||||| ||||||| |  |  ||
     MuRRS   CtgGtgGcaTatTtgTctAagAagCtgGacCcaGtgGccAgtGgaTggCccAcaCgtTta
                3920      3930      3940      3950      3960      3970
             |  ||||| || ||||| |  ||  | ||||||||||||||||| |||||  |  | |||
    MuERVC   CctGtgGccTacTtgTc.AtaAaaTtaGacCcaGtgGccAgtGggTggCcaTctTgcTta
          2710      2720       2730      2740      2750      2760


            3830      3840      3850      3860      3870      3880
     MoMLV   CggAtgGtaGcaGccAttGccGtaCtgAcaAagGatGcaGgcAagCtaAccAtgGgaCag
                  ||| |||||  | ||  | ||    || || || |  ||| ||||| |||||||
     GALV    AaaGcgGttGcaGcaGtaGcaCtcCttCtcAaaGacGctGatAagTtaAccTtgGgaCaa
               3780      3790      3800      3810      3820      3830
             |||||  |  | |||||||| || ||| | |||||  |||  ||| | ||  | || ||
     MuRVY   AaaGccAtcActGcaGtaGccCtaCttGttAaaGatActGgcAagCttActCtaGgtCag
                4240      4250      4260      4270      4280      4290
             || ||| |  || ||||||||||||| ||||| ||| ||| |||  | ||||| || |||
     MuRRS   AagGccGttGctAcaGtaGccCtaCtcGttAagGatGctGacAaaTtaActCtgGgaCag
                3980      3990      4000      4010      4020      4030
             || ||  |  || | || || |||||  | || |||||||||||| | ||||| || ||
    MuERVC   AaaGctAtcActGctGtgGcaCtaCttCtcAaaGatGctGacAaaCtgActCtaGgtCaa
           2770      2780      2790      2800      2810      2820


            3890      3900      3910      3920      3930      3940
     MoMLV   CcaCtaGtcAttCtgGccCccCatGcaGtaGagGcaCtaGtcAaaCaaCccCccGacCgc
                 |     |  | ||  |||||    | ||     | ||    ||||||||||||||
     GALV    AatGtgActGtgAttGctTccCatAgcCtcGaaAgcAtcGtgCggCaaCccCccGacCgg
               3840      3850      3860      3870      3880      3890
              |||||| ||  ||  || |||||    |  | || || ||  | || ||||| ||||
     MuRVY   CatGtgAgtG.aAtcActCccCatGcgTtaAagAgtAttGtaTGACagCccCctGacCat
                4300       4310      4320      4330      4340      4350
             ||  | | || | |  | |||||  |  |  | || ||| |  | |||||| | ||||
     MuRRS   CaaAtaActGtaGtaGccCccCacTctCttGaaAgcAttAtcCgtCagCccTcaGacCga
                4040      4050      4060      4070      4080      4090
             | ||  |  ||||||| |||| |  |  | ||||||||| |  | || ||  | || |
    MuERVC   CgaActAtaGtaGtaGacCcc.atGccAtaGaaAgcAttGttAgaCaaCcaCctGatCac
           2830      2840       2850      2860      2870      2880


            3950      3960      3970      3980      3990      4000
     MoMLV   TggCttTccAacGccCggAtgActCacTatCagGccTtgCttTtgGacAcgGacCggGtc
             ||| |  |||| ||| | |||||||| || |||  | |||| ||    |  ||  ||||
     GALV    TggAtgAccAatGccAgaAtgActCatTacCagAgcCtgCtgTta...AatGaaAggGta
               3900      3910      3920      3930         3940      3950
             |||||||| |||||| | ||||| || |||||||||||| || ||   |||||  | ||
     MuRVY   TggAtgAcaAatGccCggAtgAccCacTacCagAgcCtgTtgCta...AatGacCgaGtg
                 4360      4370      4380      4390         4400
             |||||||||||||||  |||||| ||||||||||| ||||||||    | |||  | ||
     MuRRS   TggAtgAcaAatGccTAGAtgActCacTacCagAgtCtgTtgCtg...ActGaaTggGtc
                4100      4110      4120      4130         4140      4150
             |||||||| |||||    ||||  || || ||||| || || |||   |||||  | ||
    MuERVC   TggAtgActAatGctCgcAtgAtcCatTatCagAgcCtcTttCtg...ActGagAgaGtg
            2890      2900      2910      2920      2930         2940


            4010      4020      4030      4040      4050      4060
     MoMLV   CagTtcGgaCcgGtgGtaGccCtgAacCcgGctAcgCtgCtcCcaCtgCctGagGaa...
               ||| |  ||    |  | ||| ||||| ||||| || || ||| |      ||||
     GALV    TcgTttGcgCccCctGctGtcCtaAacCcaGctAccCtaCttCcaGtcGagTcgGaaGcc
                  3960      3970      3980      3990      4000      4010
              | ||||| |  |||||  |||  || ||  | ||||| || ||      |   |   |
     MuRVY   ActTttGccCtaCctGccAtcCacAatCctAccAccCtcCtcCccAagAcgGatGccTct
          4410      4420      4430      4440      4450      4460
                ||||  |  ||||| || | ||| ||| ||
     MuRRS   GtgTttGttCctCctGctAttCtcAacCctGcc...........................
                   4160      4170      4180
               ||||   || |||||| | || || |||||
    MuERVC   CagTttAcaCccCctGctGtcCtaAatCctGctAccTtgCtgCcaTAAGtcTacGacTcc
               2950      2960      2970      2980      2990      3000


               4070      4080      4090      4100      4110      4120
     MoMLV   GggCtgCaaCacAacTgcCttGatAtcCtgGccGaaGccCacGgaAccCgaCccGacCta
                |     ||||  |||   || ||||| |||||||     ||||| ||||  ||||||
     GALV    AccCcaGtgCacAggTgcTcaGaaAtcCtcGccGaaGaaActGgaActCgaCgaGacCta
                  4020      4030      4040      4050      4060      4070
             |||   || |||   ||  | | |  ||| || |||||||||||| |  ||   || ||
     MuRVY   AccTggGtaCacCaaTgtGctGtaTacCtgGctGaaGaaActGgaTcgAgaAagGatCtg
          4470      4480      4490      4500      4510      4520
                                                           ||  ||  |||||
     MuRRS   ..............................................cgCaaAgtGatCta
                                                              4190
                                                           |    |  || ||
    MuERVC   CtaCcaGtaCacCagTgtAtgGacAtcCtgGctGaaGaaAccAgaActAggAaaGacCtc
               3010      3020      3030      3040      3050      3060


               4130      4140      4150      4160      4170      4180
     MoMLV   AcgGacCagCcgCtcCcaGacGccGacCacAccTggTacAcgGatGgaAgcAgtCtcTta
                ||||| ||  | || |     |  |  |||||||| || || || |||||| || |
     GALV    GaaGacCaaCcaTtgCccGgg...GtgCcaAccTggTatAcaGacGgtAgcAgtTtcAtc
                  4080      4090         4100      4110      4120
                |||| | | | ||| ||    |||||   |||||| ||||| ||||| || ||  |
     MuRVY   ActGacCgaActTggCcaGgt...GtgCctGacTggTacAcaGatGgtAgaAgcTttGtg
          4530      4540      4550         4560      4570      4580
               |||||   | |||||||||   || ||  ||||||||||||| || || || ||  |
     MuRRS   CgtGacCagCcgTggCcaGgt...GttCcaAacTggTacAcaGacGgaAgcAgtTtcCta
            4200      4210         4220      4230      4240      4250
                | |||||| ||||||||       || ||||||||||||||  | ||||| ||  |
    MuERVC   AcaGgcCagCcaTggCcaGgc...TgcCccAacTggTacAcaGatAgtAgcAgcTttTtg
               3070      3080         3090      3100      3110      3120


               4190      4200      4210      4220      4230      4240
     MoMLV   CaaGagGgaCagCgtAagGcgGgaGctGcgGtgAccAccGagAccGagGtaAtcTggGct
                || ||  | || |  || || ||  || |       |  |    |    | ||||||
     GALV    AcgGaaGgtAaaCggAgaGcaGggGccCcgAtcGtaGatGgcAagCggAcgGtaTggGct
           4130      4140      4150      4160      4170      4180
               ||| || |||   |||||||| |   |  | || ||  | || |      | |||||
     MuRVY   GtgGagGgaAaaTAAAgaGcaGgaGagGcaGtgGttGacAgaAaaCatGtaAttTggGca
             4590      4600      4610      4620      4630      4640
             |||
     MuRRS   GtgA........................................................
         
             ||
    MuERVC   GttGaaAgaAagTGAAaaGcaGggGcaGcaGtgGtcAacGgaAaaCaaActAtcTggGcc
                  3130      3140      3150      3160      3170      3180


               4250      4260      4270      4280      4290      4300
     MoMLV   AaaGccCtgCcaGccGggAcaTccGctCagCggGctGaaCtgAtaGcaCtcAccCagGcc
             |    ||||||||  || || || || |||  |||||||||  ||||  | || |||||
     GALV    AgcAgcCtgCcaGaaGgtAcgTcaGccCagAagGctGaaCtaGtaGccTtgAcgCagGca
           4190      4200      4210      4220      4230      4240
             ||    || |||||||| || || || ||||||||||| ||  |  |  | ||  | ||
     MuRVY   AgtTAACttCcaGaaGgaActTcgGctCagAagGctGagCtcCtgAcgCtaActAaaGct
             4650      4660      4670      4680      4690      4700

     MuRRS   ............................................................
         

    MuERVC   AgcAgcTtgCctGaaGggAcgAcaGccCagAgaGcaGagCtaAtcAcaCtcAccCaaGcc
                  3190      3200      3210      3220      3230      3240


               4310         4320      4330      4340      4350      4360
     MoMLV   CtaAagAtgGca...GaaGgtAagAagCtaAatGttTatActGatAgcCgtTatGctTtt
              ||    ||||    ||||| || ||  | ||  | || || || ||| | |||||||||
     GALV    TtaCgcCtgGcc...GaaGgaAaaAacAtcAacAtcTacAcgGacAgcAggTatGctTtt
           4250      4260         4270      4280      4290      4300
              | |  ||||||   ||||| ||    ||||| || ||||| ||||||   |||||||||
     MuRVY   CtgCagCtgGccA..GaaGgtAagGcaAtcAatAttTacActGacAgcCacTatGctTtt
             4710        4720      4730      4740      4750      4760

     MuRRS   ............................................................
         

    MuERVC   TtgCggCtgGca...Gaa...GagAacAttAatAtcTacAcaGacAgcAggCatGctTtc
                  3250            3260      3270      3280      3290


                  4370      4380      4390      4400      4410      4420
     MoMLV   GctActGccCatAtcCatGgaGaaAtaTacAgaAggCgtGggTtgCtcAcaTcaGaaGgc
             || ||||| ||||| ||||| | |||||| |    | | ||| ||||||| || |  ||
     GALV    GccActGctCatAttCatGggGcaAtaTatAagCagAggGggCtgCtcActTctGctGga
              4310      4320      4330      4340      4350      4360
             ||||| || || ||||||| ||| || ||||  || |||||||| || ||||| || ||
     MuRVY   GccAcaGccCacAttCatGagGccAtcTatAgaCaaAggGggCtaCttActTcgGcaGgc
               4770      4780      4790      4800      4810      4820

     MuRRS   ............................................................
         

    MuERVC   GccAccGctCatAtaCatGggGcaA...........AggGggTtaTtcAcaTctGctGgg
              3300      3310      3320                 3330      3340


                  4430      4440      4450      4460      4470      4480
     MoMLV   AaaGagAtcAaaAatAaaGacGagAtcTtgGccCtaCtaAaaGccCtcTttCtgCccAaa
             ||||| |||||||| ||||| || || |||||||| ||| | ||| ||  ||| || |
     GALV    AaaGatAtcAaaAacAaaGagGaaAttTtgGccCtgCtaGagGccAtcCatCtcCctAgg
              4370      4380      4390      4400      4410      4420
             || || || ||||| ||||| ||||| |||||||| |||||||| || ||  | || | |
     MuRVY   AagGaaAttAaaAatAaaGaaGaaAtcTtgGccCtcCtaGagGctAtaCacAtgCccAag
               4830      4840      4850      4860      4870      4880

     MuRRS   ............................................................
         

    MuERVC   AaaGatAttAaaAgtAaaGaaGaaAtcTtgAgcCtaTtaGaaGtcAtaCacTtgAc.Aaa
               3350      3360      3370      3380      3390      3400


                  4490      4500      4510      4520      4530      4540
     MoMLV   AgaCttAgcAtaAtcCatTgtCcaGgaCatCaaAagGgaCacAgcGccGagGctAgaGgc
              |  |   ||| ||||| ||||| || || || | ||||   | | | | ||| |  ||
     GALV    CggGtcGccAttAtcCacTgtCctGgcCacCagAggGgaAgtAacCctGtgGccActGgg
              4430      4440      4450      4460      4470      4480
                || ||||| |||| ||| | |||||| || | |||        |||| |||   ||
     MuRVY   AaaGtgGccAtcAtcCtcTgcCttGgcCatCaaAagGggCaaGgtGctGtaGccCagGga
               4890      4900      4910      4920      4930      4940

     MuRRS   ............................................................
         

    MuERVC   AacCtgGccAtcAtaCatTgcCctGgtCacCagAaaGgcCatGgtGccAtaGcaAagGga
                3410      3420      3430      3440      3450      3460


                  4550      4560      4570      4580      4590      4600
     MoMLV   AacCggAtgGctGacCaaGcgGccCgaAagGcaGccAtcAcaGagActCcaGacAccTct
             ||||||| ||| ||| | || ||     | || ||| |                   ||
     GALV    AacCggAggGccGacGagGctGcaAagCaaGccGccCtg..................Tcg
              4490      4500      4510      4520
             || | || ||| |    ||  || || || |  |  |                    |
     MuRVY   AatCagAtgGcaGgtTggGtgGccAaaCagGtgGtaCaaGgcAtcCctGcgCtg.ccTgt
               4950      4960      4970      4980      4990       5000

     MuRRS   ............................................................
         

    MuERVC   AatCagGtgGctGacCtgGctGccAcaCagGcaGccCaaGgaGctAtgAtcCtcAtaGct
                3470      3480      3490      3500      3510      3520
                  End region used for RNAse-----><-Align unreliable--...

                  4610      4620      4630               4640      4650
     MoMLV   AccCtcCtcAtaGaaAatTcaTcaCccTac.........AccTcaGaaCatTttCatTac
             |||    |  | | |    |  ||
     GALV    AccAgaGtgCtgGcaGgaActAca....................................
            4530      4540      4550
             |         | | || | | | |
     MuRVY   AgaGag...AtaGaaGaaCcaAaaGagAcc.........CcaGagAtcAgtTttGa.Tac
                   5010      5020      5030               5040       5050

     MuRRS   ............................................................
         

    MuERVC   AgaGagAaaAatAaaGaaCttAagGatCatTatGaaCtcAaaGccAcaGgcTttGacTac
                3530      3540      3550      3560      3570      3580


                 4660      4670               4680      4690
     MoMLV   AcaGtgActGatAtaAagGac.........CtaAccAagTtgGggGccAtt......Tat
                            ||             | |       | | | ||  |       |
     GALV    ...............AaaCct.........CaaGagCcaAtcGagCccGct......Caa
                                          4560      4570            4580
                             |             |  ||   |   | |    |
     MuRVY   AccTccAaaGatTatGagAtgAatGaaGaaCttGaaGaaCaaGggAagTccAaaGgaTtt
                   5060      5070      5080      5090      5100      5110

     MuRRS   ............................................................
         

    MuERVC   ActCcaGaaGatCagGaaCtaAcaGgcAaaAtaCctGagAtaTgt...Agg......Tat
                3590      3600      3610      3620         3630


            4700      4710      4720      4730      4740      4750
     MoMLV   GatAaaAcaAagAagTatTggGtcTacCaaGgaAaaCctGtgAtgCctGacCagTttAct
             || || || | |  |                  |      | |  ||||||| |
     GALV    GaaAagAccAggCcg..................AggGagCtcAccCctGacCgg...Gga
                  4590                        4600      4610         4620
                 |     ||                     |      |    ||  |  |    |||
     MuRVY   AgcTacCatGggAtaGcaAgaAcaAaaGaaGg.AaaAttAttTtgCccTatAgaGaaGga
                   5120      5130      5140       5150      5160

     MuRRS   ............................................................
         

    MuERVC   ActCccCagGaaGtgGctAagAccCaaGatGgtTgcTtgGtcCtgCccAccAaaGaaGga
               3640      3650      3660      3670      3680      3690


            4760      4770      4780      4790      4800      4810
     MoMLV   TttGaaTtaTtaGacTttCttCatCagCtgActCacCtcAgcTtcTcaAaaAtgAagGct
                |||||  |  |     | |||||| | |||||| |  |      |||  |
     GALV    AaaGaaTtcAttAagCggTtaCatCagTtaActCacTtaGgaCcaGaaAagCttCtcCaa
                   4630      4640      4650      4660      4670      4680
             || | ||   | || ||  | ||||   | || ||  | ||| || | ||  |     |
     MuRVY   AagGcaTatGtaAaaCgcAtcCatCccCtgAcaCatCtgGgaGcaAagAaaTtgTgtGat
          5170      5180      5190      5200      5210      5220

     MuRRS   ............................................................
         

    MuERVC   CaaCaaTatGtaGccAatGtaCacCagCtcAtgCacTtaGaaActAagAatCtcAaaGaa
               3700      3710      3720      3730      3740      3750


            4820      4830      4840      4850      4860      4870
     MoMLV   CtcCtaGagAgaAgcCacAgtCccTacTacAtgCtgAacCggGatCgaAcaCtcAaaAat
             ||  |  |  |       |    |  |  ||| | |||||   |     || |     |
     GALV    CtaGtgAacCgt......AccAgcCtcCtcAtcCcgAacCtcCaaTctGcaGttCgcGaa
                   4690            4700      4710      4720      4730
               | |  |           | | |       | |  |  || | |             ||
     MuRVY   TAAAtcCagTcc......TcaAacTatTatGtgCtcAggCtaCtaGac......GaaCaa
          5230      5240            5250      5260      5270

     MuRRS   ............................................................
         

    MuERVC   AttGttAagAac......TcaAacTacTatGttAtaAaaCtgTtcAacGttGccCagGag
               3760            3770      3780      3790      3800


            4880      4890      4900      4910      4920      4930
     MoMLV   AtcActGagAccTgcAaaGctTgtGcaCaaGtcAacGccAgcAagTctGccGttAaaCag
              ||||       ||  | ||||||||       || ||   ||   |   |    |
     GALV    GtcAccAgtCagTgtCagGctTgtGccAtgActAatGcgGtcAccAccTacAgaGagAcc
               4740      4750      4760      4770      4780      4790
             || || |     ||  | |  ||| || | | |||||| |    | |||  |  ||
     MuRVY   GtaActAaaTccTgc.aaGtgTgtTccCtaAttAatGctGcaCgcCccTttAatGaaCaa
            5280      5290       5300      5310      5320      5330

     MuRRS   ............................................................
         

    MuERVC   AttGttAatAaaTgcCaaGccTgtGccCtcAccAatGccAggCacCacAagAacGtcCcc
           3810      3820      3830      3840      3850      3860


            4940      4950      4960      4970      4980      4990
     MoMLV   GgaActAggGtcCgcGggCatCggCccGgcActCatTggGagAtcGatTtcAccGagAta
             ||||  |||   || ||  |||| ||||||    | |||||| | || ||||| || |||
     GALV    GgaAaaAggCaaCgaGgaGatCgaCccGgcGtgTacTggGagGtaGacTtcAcaGaaAta
               4800      4810      4820      4830      4840      4850
             |||||| | ||||  || ||| |||| || || |||||||| ||||||||||| || ||
     MuRVY   GgaAaaCgaCaaCtgGggGatAgaCctGgaGttTacTggGaaGtaGacTtcActGagAtc
             5340      5350      5360      5370      5380      5390

     MuRRS   ............................................................
         

    MuERVC   GgaAggTggCtaAgaGgtGacCggCcaGggGcaTacTGAAaaGtaAatTttActGaaGtt
           3870      3880      3890      3900      3910      3920
                            <-- begin region used as Int domain

            5000      5010      5020      5030      5040      5050
     MoMLV   AagCccGgaTtgTatGgcTatAaaTatCttCtaGttTttAtaGatAccTttTctGgcTgg
             ||||| ||   ||||||  | || |||||  |||| || |||||||| || || || |||
     GALV    AagCctGgtCggTatGgaAacAagTatCtgTtaGtaTtcAtaGatActTtcTccGgaTgg
               4860      4870      4880      4890      4900      4910
             || || ||    ||||| || || ||  |  |||| |||||||| |||||||| ||||||
     MuRVY   AaaCcaGgaAaaTatGgtAatAaaTAGTtaCtaGttTtcAtaGacActTtcTcaGgaTgg
             5400      5410      5420      5430      5440      5450
                                             ||||||  |||| || || |||||||||
     MuRRS   ...............................aaGttTttGtaGatAccTttTcaGgaTgg
                                          4260      4270      4280
                                             ||||||| |||| || |||| ||| |||
    MuERVC   AaaCcaGccAagTacGgtAatAaaTatCtaTtaGttTttAtaGacActTttTGAGggTgg
           3930      3940      3950      3960      3970      3980


            5060      5070      5080      5090      5100      5110
     MoMLV   AtaGaaGccTtcCcaAccAagAaaGaaAccGccAagGtcGtaAccAagAagCtaCtaGag
              ||||||| || || ||||| |  ||||| |||    ||||    || ||  || ||||
     GALV    GtaGaaGctTttCctAccAaaActGaaAcgGccCtaAtcGtcTgtAaaAaaAtaTtaGaa
               4920      4930      4940      4950      4960      4970
             |||||||| || || |||||  ||||||| || |   | ||    || || ||  | |||
     MuRVY   GtaGaaGccTtcCccAccAagTctGaaActGctCagGtgGtgAccAagAagAtcGttGaa
             5460      5470      5480      5490      5500      5510
             || || |||| | | || ||    ||||||||  | || |||   || || ||| |  ||
     MuRRS   GtgGagGccTacAcaAcaAa.AaaGaaActGccAatGttGtgTttAaaAaaAtcCtaAaa
            4290      4300       4310      4320      4330      4340
                                     || || ||||||||||||      || ||  || |
    MuERVC   ........................GagAccGccAatGttGtgAc...gAagAtaTtaGag
                                   3990      4000         4010      4020


            5120      5130      5140      5150      5160      5170
     MoMLV   GagAtcTtcCccAggTtcGgcAtgCctCagGtaTtgGgaActGacAatGggCctGccTtc
             || ||  | ||| | ||||| || ||| ||||| | ||  | |||||||| || |||||
     GALV    GaaAttCtaCccCgcTtcGggAtcCctAagGtaCtcGggTcaGacAatGgcCcgGccTtt
               4980      4990      5000      5010      5020      5030
             || || |||||||  || |||||||| |  ||| |  | |  || ||||| || || |||
     MuRVY   GagAtcCtaCccCagTttGggAtcCccAgtGtaAtaTgcTtgGaaAatGggCctGcaTtt
             5520      5530      5540      5550      5560      5570
              | ||  | || | |||| | || || |  |||||| | |  || ||||  ||  | | |
     MuRRS   AaaAttTttCctCggTttAgaAtaCctAagGtaAtaGggTctGacAatGaaCccAccTct
             4350      4360      4370      4380      4390      4400
              ||||||| ||  | ||| | |  ||||||||| |    || |||||||  |||  ||
    MuERVC   GaaAttTtcCcaAgaTttGgcA.cCctAagGtaCttTcaTcaGacAatGgcCcc.tcTtc
                  4030      4040       4050      4060      4070


            5180      5190      5200      5210      5220      5230
     MoMLV   GtcTccAagGtgAgtCagAcaGtgGccGatCtgTtgGggAttGatTggAaaTtaCatTgt
             ||  |  |||| ||||||  | |||||  ||   |||||||  ||||||| |||||||||
     GALV    GttGctCagGtaAgtCagGgaCtgGccActCaaCtgGggAtaAatTggAagTtaCatTgt
               5040      5050      5060      5070      5080      5090
             ||||| ||||||||||||| | | ||||| ||  ||||||  |||||||||||| |||||
     MuRVY   GttGccCagGtaAgtCagGaaTtaGccAcaCatTtgGggAacAatTggAagTtaTatTgt
             5580      5590      5600      5610      5620      5630
             ||||||||||||||||||| | | ||||  || |||||||   ||| ||| ||| |||||
     MuRRS   GttGccCagGtaAgtCagGgaCtgGccAtcCaaTtgGggAttGatTAGAaaTtaCatTgt
             4410      4420      4430      4440      4450      4460
             ||||||||||||||||| |   | ||||  |||||||||||||||| ||| |||||||||
    MuERVC   GttGccCagGtaAgtCaaGcgTtaGccAgaCaaTtgGggAttGatTggAagTtaCatTgt
          4080      4090      4100      4110      4120      4130


            5240      5250      5260      5270      5280      5290
     MoMLV   GcaTacAgaCccCaaAgcTcaGgcCagGtaGaaAgaAtgAatAgaAccAtcAagGagAct
             || || |||||||| |||||||| ||||||||||||||||| ||||| || || |||||
     GALV    GcgTatAgaCccCagAgcTcaGgtCagGtaGaaAgaAtgAacAgaAcaAttAaaGagAcc
               5100      5110      5120      5130      5140      5150
             || |||| ||||||||| ||||   |||||||| |||||||||| ||  | ||||| |||
     MuRVY   GctTatAaaCccCagAgaTcaGtaGagGtaGaaGgaAtgAacAggActCtaAaaGaaAcc
             5640      5650      5660      5670      5680      5690
             | ||||| ||||||||| ||||   |||||||| |        | |    |||| ||| |
     MuRRS   GttTatAgaCccCagAgtTcaGgcTAGGtaGaaAg........gaAaaAaaAaaAaaAac
             4470      4480      4490      4500              4510
             ||||||||| ||||||||||| |  || ||||  |        ||| |  |||| |||
    MuERVC   GttTatAgaAccCagAgtTcaAgaCagTtaGagCggAtaAatAgaAcaTtaAaaGaaAca
          4140      4150      4160      4170      4180      4190


            5300      5310      5320      5330      5340      5350
     MoMLV   TtaActAaaTtaAcgCttGcaActGgcTctAgaGacTggGtgCtcCtaCtcCccTtaGcc
             || || |||||| |  | |  || ||    | ||||||||||  ||| || ||||||||
     GALV    TtgAccAaaTtaGccTtaGagAccGgtGgaAaaGacTggGtgAccCtcCttCccTtaGcg
               5160      5170      5180      5190      5200      5210
              | || |||||||| || |||||| |||| |||||||||||||||||||||||||| | |
     MuRVY   CtaActAaaTtaGctTttGagAccAgtGgtAaaGacTggGtgAccCtcCttCccTttGtg
             5700      5710      5720      5730      5740      5750
              | |||||||||   || |||||| |  | |   |||||    |||||||||||||||
     MuRRS   TtgActAaaTtaAtcTtaGagAccGgcAgaAgtTacTggAcaGccCtcCttCccTttGct
           4520      4530      4540      4550      4560      4570
             || ||||||||    |||||||| ||| |||   | ||| |||| |||||| ||||  |
    MuERVC   TtaActAaaTtgGctTtaGagActGgcTGAAaaGatTggGcaGctCtcCttTccTtcCcc
          4200      4210      4220      4230      4240      4250


            5360      5370      5380      5390      5400      5410
     MoMLV   CtgTacCgaGccCgcAacAcgCcgGgcCccCatGgcCtcAccCcaTatGagAtcTtaTat
             |||    | ||| | || || || ||||    |||  | || || ||||| ||  | |||
     GALV    CtgCttAggGccAggAatAccCctGgcCggTttGgtTtaActCctTatGaaAttCtcTat
               5220      5230      5240      5250      5260      5270
             || ||||||||| | || || |||||  | || ||  | |||||||||||||||||  ||
     MuRVY   CtcCttAggGccCgaAacAcaCctGggTgtTtcGgcAttActCctTatGaaAttCtaCat
             5760      5770      5780      5790      5800      5810
              |  |  | |   | |||||||| ||    ||     ||||||| ||| |||| ||  ||
     MuRRS   TtgTtcCgtGttAggAacAcaCcaGgaAaaTttAagCttActCcaTatAaaAtcCtgTat
           4580      4590      4600      4610      4620      4630
             |||||| | ||   |||||| ||||| |||||||| ||||  || | | || ||||  ||
    MuERVC   TtgTtcTGAGtaCagAacAccCcaGggAaaTttAaaCttAaaCccTttGaaCtcCtcCat
          4260      4270      4280      4290      4300      4310


            5420      5430      5440      5450      5460            5470
     MoMLV   GggGcaCccCcgCccCttGtaAacTtcCctGacCctGacAtgAcaAga......GttAct
             || | ||| || ||| |  |  | |     ||  ||       |          | ||
     GALV    GgaGgaCcaCccCccAtaCttGagTctGgaGaaActTtgGgtCccGat......GatAga
               5280      5290      5300      5310      5320
             || |||||||||||| |    |||||||  || |  |||  | ||  |        |
     MuRVY   GggGgaCcaCccCccTtgAccGagTctGatGagAtaTtgTatTccAgtGctGacTctTcc
             5820      5830      5840      5850      5860      5870
             |||||     | |||  |  |      |   ||| | |||       |    | | |  |
     MuRRS   GggGggGggGctCccCcgCtcActAaaGtaAagAgaGtgTtaGatCctTtaAaaTttAac
           4640      4650      4660      4670      4680      4690
             |||||     | |||           ||     |   |      |  |       ||
    MuERVC   GggGgaCccCccCccTtaAcaGaaGcaGggGgaAtgTttGacCctGat......GttCct
          4320      4330      4340      4350      4360            4370


                  5480      5490      5500      5510      5520      5530
     MoMLV   AacAgcCccTctCtcCaaGctCacTtaCagGctCtcTacTtaGtcCagCacGaaGtcTgg
                  |||     |     |||||||| ||||| |  |  | ||   |  | || |||||
     GALV    TttCtcCctGtcTtaTttActCacTtaAagGctTtaGaaAttGtaAggAccCaaAtcTgg
           5330      5340      5350      5360      5370      5380
             |  | | ||    | ||||| || |||||||| |||||| |||| || || || ||||||
     MuRVY   TcaCccTctTctCtgTttAccCatTtaAagGccTtaGaaGttGtcAgaAcaCagAtcTgg
             5880      5890      5900      5910      5920      5930
              |     ||||  |||||||     |||| ||| ||||||| |||||||       |||
     MuRRS   CctGgtCctTcaTtgTttAcaTgcAtaAaaGccCtaGaaGtgGtcAgaAacAccGccTGA
           4700      4710      4720      4730      4740      4750
              ||   |   |  || |  |  || | || ||||||||| |||||| |||    ||||||
    MuERVC   TctTccCaaCccCtgCtaGctCgcTtgAagGccCtaGaaCtgGtcAaaAaaGatGccTGA
                4380      4390      4400      4410      4420      4430


                  5540      5550      5560      5570      5580      5590
     MoMLV   AgaCctCtgGcgGcaGccTacCaaGaaCaaCtgGacCgaCcgGtgGtaCctCacCctTac
                |   |    |  |  ||  |          |            | |||||||| | |
     GALV    GacCagAtcAaaGagGtgTatAagCct......GgtAccGtaAcaAtcCctCacCcgTtc
           5390      5400      5410            5420      5430      5440
             |||||||| ||||| |  || |  ||       || | |    || | || |||  ||||
     MuRVY   GacCagAttAaaGaaGcaTacAcaCca......GggAgcAgtGcaGtgCccCacAggTtc
             5940      5950      5960            5970      5980      5990
             || ||| | ||||| || |||  ||||      ||   |       | || |||  ||||
     MuRRS   GagCagCtcAaaGagGccTacGaaCca......GgaGacTtgAtgAtaCcaCacCagTtc
           4760      4770      4780            4790      4800      4810
             ||   ||| ||| |    || ||||||       ||  |       |  |||   | ||
    MuERVC   GaaTtgCtgAaa.aaAagTatGaaCca......AgaAccAccGcaGtgTcaCgtAaaTtt
                4440       4450            4460      4470      4480


                  5600      5610      5620      5630      5640      5650
     MoMLV   CgaGtcGgcGacAcaGtgTggGtcCgcCgaCacCagActAagAacCtaGaaCctCgcTgg
             |  ||||| ||   ||||   ||| | || || |   | |  | ||| || ||||| |||
     GALV    CagGtcGggGatCaaGtgCttGtcAgaCgcCatCgaCccAgcAgcCttGagCctCggTgg
                 5450      5460      5470      5480      5490      5500
             |||||||| ||   ||| || ||| |||  |||||| |  |||  | |||||| ||||||
     MuRVY   CagGtcGg.GacTcaGttCtgGtcTGACaaCatCgaGctGgcAtgCatGagCcaCggTgg
                    6000      6010      6020      6030      6040
             ||||| || ||  ||||||| ||||||| |||||||||||  |  |   | ||  | |
     MuRRS   CagGtgGgaGatGcaGttCtcGtcTGACgaCatCgaGctGagAatCtgAaaCctTgtTAA
                 4820      4830      4840      4850      4860      4870
             ||  |  | ||| | || || ||||  || ||  |  |    || ||  ||||    |
    MuERVC   CaaAtaAggGatCctGtcCtgGtcTAGCgcCacTgtTccAgcAacCttGaaCccAaaTgg
             4490      4500      4510      4520      4530      4540


                  5660      5670      5680      5690      5700      5710
     MoMLV   AaaGgaCctTacAcaGtcCtgCtgAccAccCccAccGccCtcAaaGtaGacGgcAtcGca
             ||||| || |||   ||  ||||||| ||||| |||||  | ||||| || || || ||
     GALV    AaaGgcCcaTacCtgGtgTtgCtgActAccCcgAccGcgGtaAaaGtcGatGgtAttGct
                                                              Met for env
                 5510      5520      5530      5540      5550      5560
             || || || || || ||| ||||||| | | |    |  | ||| || ||  | || |||
     MuRVY   AagGgaCctTatCtaGtgCtgCtgAccAgcGccGagGtcGcaAagGtaGacAggAtcGct
          6050      6060      6070      6080      6090      6100
             || ||  |||   | ||||| ||||||| | ||   |  |  || | |||  | |||
     MuRRS   AaaGgcTctTgcAtcGtgCtaCtgAccAccCccActGctGtcAaaGaaGaaGgcAtcTac
                 4880      4890      4900      4910      4920      4930
             |||||  | | | | ||||| |||||
    MuERVC   AaaGgaCccTacCtgGtgCttCtgAc.....remainder MuERVC pol deleted 
             4550      4560      4570
               Used as Int    ---------><--  alignment unreliable--...
                                           Trying to align the end of pol
                                           or the beginning of env which
                                           overlaps in a different frame
                                           tends to produce conflicting
                                           alignments, neither of which
                                           is convincing.

                  5720         5730      5740         5750      5760
     MoMLV   GctTggAta...CacGccGccCacGtgAagGc...tGccGacCccGggGgtGgaCcaTcc
             || ||| |    || ||  | ||| | ||  |   |||    ||   ||    |
     GALV    GccTggGtc...CatGctTctCacCtcAaaCc...tGcaCcaCctTcgGcaCcaGatGag
                          Met (internal): better candidate for
                          homologue to initiator ATGs of env of
                          other viruses.

                 5570         5580      5590         5600      5610
             |||||| ||   || || || ||| | |||      |||||| |   ||    ||| ||
     MuRVY   GccTggAtc...CacGcaTccCacAtaAaaAa.AagGcaCcaAccA.gGatGaaGacGac
                                                              Met for env
                                                  possible pol ter

          6110         6120      6130       6140      6150       6160
             || ||  |    |||| |   |
     MuRRS   GcaTgcCttT..CacGtaAagC  remainder MuRRS pol deleted         
                 4940        4950



              5770      5780      5790      5800      5810      5820
     MoMLV   TctAgaCtgAcaTggCgcGttCaaCgcTctCaaAacCccTtaAaaAtaAggTtaAccCgc
                        Met for env
             ||          |||    |  ||
     GALV    Tcc.........TggGagCtgGaaAagA................................
                      5620      5630
               |         |||    |||     |
     MuRVY   CtcAgaGaaAatTggAcaGtgGccGctA................................
               6170      6180      6190
         


              5830      5840      5850      5860      5870      5880
     MoMLV   GagGccCccTAAtccccttaattcttctgatgctcagaggggtcagtactgcttcgcccg
                     ||  | |  |        |  || | || | | | ||   |  ||  ||
     GALV    ........ctGatCatCctCttAagCtgCgtAttCggCggCggCggGacGagTctGcaAa
                      5640      5650      5660      5670      5680
                     ||||| || |||  |   ||| |           || | ||
     MuRVY   ........ctGatAatTctCaaActGtgCctT.........cgCtgTacAccAacCttGg
                         6200      6210               6220      6230
                     <-- Begin Env conserved seq. in overlapping frame.
                     GALV and MoMLV frameshift relative to one another
                     in C terminal of pol and finish at different 
                     stop codons.


              5890    
     MoMLV   gctcc
                    
     GALV    aTAA
            5690 
              |||
     MuRVY   gTAA
              6240